|
Journal Number 96
August 2005
THE COLUMN
Caladenia Identification from Marginal Calli
By Eric Scanlen
| |
|
|

|
|
ZERO TO ONE MARGINAL CALLUS
C. alata R. Br. has one golden topped, blade-like callus, reduced at times to a solitary green or golden stump
(cover J94) or none at all.
Golden midlobe tip curled under, acute, white to pink tepals,
solitary flowers open 6 Sept to 24 Oct at Te Paki, mid October
at Rainbow Mountain.
Scrubby track-sides in dappled shade, sparse.
Drawn from two slides of the same unusually late flower at the Earth Wall, Te Paki at 7 Oct 00. |
| |
|
|

|
|
ONE ONLY MARGINAL CALLUS
C. "speckles" (disputed) of Allan Ducker's with red stem, bracts and ovary, the last with 3 green stripes, one golden topped, rod-like BMC rising from the white base of the yellow, triangular midlobe with raised midrib.
Solitary pink to white flowers, dorsal sepal speckled pale plum inside, tepals acute with a broad but tapering stripe of red glands down the outer midribs. (Journal 94:35).
Column top
is dark red.
Te Paki in kanuka or clay road side. See J82:15, J72:18 Fig. 9,
flowers 25 Oct - 10 Nov.
Drawn from 2 slides of one flower on Pink Track, Te Paki from 1/11/99 |
| |
|
|

|
|
ONE TO 1½ MARGINAL CALLI
C. aff. bartlettii of Doug McCrae's has one to 1½ BMC, all-yellow and rounded, curling forward then margin bumpy to the tip of the all yellow midlobe with straight cross section to turned down margins. The ½ BMC is a big bump rather than a callus.
Note, recurved and rounded sepals, tiny, solitary, pink flowers
from Kaitaia 25 Sep to 15 Oct 1912, Sweetwater (very hairy)
26-31 Oct 1989 and Te Paki 31 Oct to 5 Nov, on scrubby track sides, rare.
See J94:35 Fig. 6, J78:20 & Plates 11,12.
Whole flower from Caladenia Track, Shenstone Block, Te Paki, 5 Nov 00. Midlobe from NZ Walkway, Scott Point 31 Oct 04. |
| |
|
|

|
|
TWO MARGINAL CALLI and rounded sepal tips
C. bartlettii (Hatch) D.L. Jones, Molloy & M.A. Clem., has two BMC as a rule, sometimes only one, or rarely two on one side and one on the other but always yellow-topped, rounded calli leaning towards the narrow, yellow, trough shaped midlobe.
Note, four tepals in a flat plane, cerise all over or cerise pink fading to white at the base.
Red (rarely green) stemmed, sparse, Te Paki 27 Sept - 5 Nov,
to Nelson in late Oct, favours damp clay soil on track sides.
See J82:15, J75:36, top right, J72:27,28.
Drawn from a rare green stemmed flower from Scott Point, 27 Oct 01, midlobe from a red stemmed specimen, 22 Oct 03, below Mangatangi Dam, Hunua's. |
| |
|
|

|
|
2½ MARGINAL CALLI sometimes only 1½ rounded sepal tips
C aff. pusilla of Ian St George's, yellow-topped and rounded BMC leaning towards the all yellow, triangular midlobe plus lateral petals turned up at the tips and the dorsal sepal always curled down over the column.
Disc calli have red legs and yellow tops, dark red bars in lower labellum wings merge to solid colour on the disc. Tepal backs liberally coated with dark red glands.
Solitary pink flowers on clay soil track sides, in a North Island disjunct distribution of Te Paki 30 Sept (1½ BMC) to 10 Oct, Kaimaumau J62:24 as C. minor, and with 2½ BMC at Awhitu 19 Oct, Te Puke to Rotorua roadside, Moki Rd Uruti 18-23 Oct, J92:13, and Puffer Track Kaitoke, J82:15, rare.
See the Puffer specimen in the 1996 Field Guide as C. minor.
Drawn from a 11 Oct 02, 2½ BMC flower from Papawiri Hill, Te Paki; midlobe from 19 Oct 04, Awhitu Central. |
| |
|
|

|
|
THREE OR FOUR MARGINAL CALLI plus acute tepals
C. "nitida rosea" of H.B. Matthews has 4, 3 or rarely 2 BMC;
1, 2
or rarely, 3 flowers opening singly; dorsal sepal inner
not always speckled but otherwise close to the smaller
C. "speckles".
White to pink, favours mature kanuka in a North Island disjunct distribution of
Te Paki (6 Oct to 6 Nov) J87:25, J77:21, as Petalochilus aff. fuscatus; J62:9, Taiharmu, Coromandel Oct 1984; Moki Rd Uruti (18 Dec 03) J92:14.
The last had atypical crimped labellum wings, acuminate tepals (not just acute) and no raised midrib to the labellum indicating possibly millennia separation from northern colonies.
Drawn from a Scott Point flower of 27 Oct 01 with 3 BMC; midlobe from a 4 BMC specimen, 18 Oct 96, Shenstone Track, Te Paki. |
| |
|
|

|
|
C. "chloroleuca" H.B. Matthews' disputed taxon, 2-5 flowered, has
3 sometimes 4, yellow BMC with only serrations to the tip of the white triangular midlobe, canoe prow sepals and greenish, labellum wings almost solid maroon inside.
At present, only at Te Paki, mid October J94:35; 72:27, and
Moki Rd, late November J92:19.
Appears to hybridise with
C. minor which has only 1 to 2
flowers and spaced red bars in the labellum wings.
Drawn from one slide of a 23 Oct 98 flower by the Shenstone Track,
Te Paki. |
| |
|
|

|
|
C. "red stem" (disputed) of Gordon Sylvester's is all maroon in bud, perhaps a tinge of olive on the tepal backs as at Nelson, has 2 to 5 flowers, off white tepals each with 3 parallel, red veins.
BMC are numerous as in C. minor with yellow expanded tops.
The first one to three BMC uniquely have maroon stems like
the disc calli.
Known from light scrub along the Puffer Track, Kaitoke (mid
December) J66:26, from the Baton Valley, Nelson and Gordon
reports it from around Wellington Harbour and spots on the
S.I. West Coast.
Drawn from 2 shots of the first flower to open on the Puffer Track,
Kaitoke colony, 6 Dec 97. |
| |
|
|

|
|
NUMEROUS (5 TO 9) MARGINAL CALLI all with acute
to minutely acuminate tepals
C. minor Hook. f., alias C. chlorostyla has 5 to 7 discernable, creamy yellow BMC, usually short and flattened sometimes stalky and bent forward but always reducing towards the tip of the long, narrowing, white to creamy midlobe.
Note, 1 to 2 flowers, palest green tepals, hairy back, to the dorsal sepal, bright green stem, bracts, leaf and column back; hence Doug McCrae's tag, C. "green column".
Thrives in kanuka or beech debris, opens 4-28 Oct at Te Paki J82:15,17, 26 Nov at Whenua Tapu, Wellington and early January at Lake Hauroko.
Spaced red bars in the labellum wings and column plus the yellow topped calli, give a break from green on this plant.
Drawn from 2 shots of one flower, 28 Oct 01, track to Radar Bush,
Te Paki. |
| |
|
|

|
|
C. aff. chlorostyla of Bruce Irwin's, was included in Doug's
C. "green column" and the Column's C. minor in J72:27,28 but its peak flowering is generally a fortnight later than C. minor, it has off white rather than green tepals and has closely packed red glands on the dorsal sepal back. Its green column is red banded inside and has two red blobs on top.
BMC are flattened, 1-3 near the base are hooked forward, stems and ovaries can be green (J75:36, Fig. 4, left) or maroon (J75:36 Fig. 5), the latter have 3 green wedges up the maroon ovaries.
Flowering mid October at Te Paki, mid December at Pukeiti, late December at St Arnaud to late January at Lake Manapouri South Arm, track sides in kanuka or beech; ref. J92:19; 88:21; 86:32.
Drawn from shots of 2 red stemmers; whole flowers at Cheesemans Track, Te Paki, 13 Oct 02; midlobe at the Brunner Peninsula, St Arnaud, 22 Dec 02 where at first it was taken to be the closely related C. "red stem". |
| |
|
|

|
|
C. nothofageti D.L. Jones, Molloy & M.A. Clem., has up to 9 BMC, some in the middle, hooked forward as in closely related C. aff. chlorostyla.
1 to 2 almost all white flowers are out in December at the Puffer Track, J66:25 and January (after C. minor has faded) at Lake Hauroko, Fiordland.
This is not an albino because it has creamy calli tops - which were very dark under ultra-violet light J86:5 - and a grass-green top to the column, J90:26 (open on 23 Jan 04) at Slab Hut near Monowai.
This is an exception to the specified "reddish barred labellum" but, some with pale pink bars were at Lake Hauroko, J91:12 on 20 Jan 04.
Flower drawn from Brunner Peninsula, say 30m from C. aff. chlorostyla on 22 Dec 02. Midlobe from a pink barred specimen at Lake Hauroko, Fiordland, 20 Jan 04. |
| |
|
|

|
|
NUMEROUS MARGINAL YELLOW BUMPS
RATHER THAN CALLI
C aff. variegata (disputed) tagged by the Column for its
regular double row of yellow disc calli (J94:36 Fig. 11) with
no stray calli.
The largest of NZ's Caladenias; single pink flowers (rarely twin) with acute tepals. BMC are either a variable array of bumps or the margin may be entire with one or two rounded notches etc. to the tip of the triangular yellow but white centred midlobe.
The midlobe margin is the clearest physical difference from the smaller C. "nitida-rosea" and
C. "speckles" above.
But colour-wise, C. aff. variegata has that almost luminous green column back, tepal backs have a central, pale green band and a narrower, broken strip of red glands down the midrib.
It favours dappled shade in Pinus radiata or beech forest from Iwitahi early Dec, J78 Plate 2, top right, to the Aorangi's in late Nov, J94:35,36 to Otago J82:15.
Drawn from a flower at Iwitahi, 13 Dec 97, midlobe from Mill Track, Aorangi's, 27 Nov 04 |
| |
|
|

|
|
C. variegata Col, an exception to the specified 2 rows of disc calli because stray calli are all round, the 2 rows may be doubled up (J72:27 Fig. 3) or have the whole disc cluttered with calli (Aorangi's, J94:36 Fig. 13), which may advance in a chevron down the top of the midlobe but not to the tip.
Otherwise close to C. aff. variegata, growing in the same areas.
At Iwitahi, one will predominate some seasons and the other
will in other seasons.
Drawn from a flower at Iwitahi, 2 Dec 94, cluttered disc calli from Sutherlands Track, Aorangi's, 27 Nov 04. |
| |
|
|
Moral? Basal marginal calli on the labellum midlobe are good, prime, identifiers of Caladenia
but other traits need roping in to be sure of which taxon or species you are looking at.
Acknowledgements : Many thanks to Doreen Abraham, Trish Aspin, Ernie Corbett, Allan Ducker, Dan Hatch,
Bruce Irwin, Brian Molloy, Gary Penniall, Graham Marshall, Phil Mitchell, Mark Moorhouse, Michael Pratt, Kelly Rennell,
Ian St George, John Smith-Dodsworth, Sid Smithies, Gordon Sylvester, Georgina Upson and any other great Caladenia
finders I might have missed.
RH & HB Matthews' and Doug McCrae's amateur but enlightened work in this field from earlier times, is also
gratefully acknowledged.
|